Mumble's son, Erik, is struggling to realize his talents in the Emperor Penguin world. Meanwhile, Mumble and his family and friends discover a new threat their home -- one that will take everyone working together to save them.
The magically long-haired Rapunzel has spent her entire life in a tower, but now that a runaway thief has stumbled upon her, she is about to discover the world for the first time, and who she really is.
Manny, Sid, and Diego discover that the Ice Age is coming to an end, and join everybody for a journey to higher ground. On the trip, they discover that Manny, in fact, is not the last of the wooly mammoths.